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RE: Re: Anthocyanin Pigment Inheritence

Thanks Chuck

This is great information

Colleen Modra
Adelaide Hills 
South Australia


-----Original Message-----
From: owner-iris@hort.net [mailto:owner-iris@hort.net] On Behalf Of
Sent: Saturday, 26 July 2008 12:17 PM
To: iris@hort.net
Subject: [iris] Re: Anthocyanin Pigment Inheritence

In response to queries re glaciatas.

There is basically three genetic lines to remove anthocyanin. That is 
three sets of genes that have been used in iris.  To produce 
anthocyanin, particularly delphinidin, there is about 22 biochemical 
steps  in the production. Blocking any of these will stop the 
production of anthocyanin.  These are called structural genes. In 
addition you can have  control genes that prevent certain genes from 
acting or by themselves  remove anathocyanin.

But lets focus on what we  know.

There is recessive reduction of anthocyain, This used to be called 
recessive white and  the symbol used was "w"  and blues were "W". But 
this is incorrect as  yellows, pinks, and oranges etc  all  can exsist 
with four sets of "w", ie w/w/w/w  genotypes.  One of this type crossed 
with a   flower with anthocyanin produces all anthocyanin flowers , as 
long as no recessives were carried by the anthocyanin flower (brown. 
ed. purple, blue etc)

There is the dominant reduction of anthocyanin , labelled "I" and we 
can use this symbol. This is the commonest gene used to produce pinks, 
yellows oranges, apricots etc.   A cross of one of these (having one 
dosage of I )  to a flower having anthocyanin (ignoring all recessives 
present) will  produce  half flowers with anthocuyanin and half without.

Not the use of the term "reduction", not removal. Multiple dosages of I 
will probaly  reduce anthocyanin even more.

Plicata genetics are complicated. Basically a plicata is a plant having 
four plicata genes,  selecting from the three known alleles. Luminata, 
glaciata and luminata.  A plicata can also have other genes such as I 
or wwww, removing anthocyanin. Thus we get a plicata such as Laced 
Cotton.  There is nothing  in definition of plicata genotype  
indicating it has to have anthocyanin or any fixed pattern of 
distribution. Known patterns include  classic plicata, luminata, 
luminata-plicata, zonal and glaciata. We don't know what the pattern is 
when  a glaciata has four luminata genes. It had been assumed for a 
long time that luminata pttern had four luminata genes but  this is not 
so.  Also any of these patterns can also have an I gene and we can get 
a plicata pattern in cartenoid pigments with  a plicata distribution. 
Now called a yellow plicata

The glaciata , which is four sets of  the plicata glaciata gene , also 
has no anthocyanin. The removal of anthocyanin this way  produces an 
exceptional clean flower.  But an exceptionally clean flower is not 
necessarily a glaciata.

A glaciata  flower crossed with a wwww  will produce all offspring with 
anthrocyanin. For example a white glaciata X  recessive white, produces 
all blue offspring.

The glaciata is a recessive of a recessive (plicata) and has absolutely 
no effect on pigment unless  paired up as one of four plicata genes. 
And only shows glaciata  when in a set of four glaciata genes. So is of 
no interest unless you are working plicatas. Of course working a line 
of glaciatas to produce  exceptionally clean non anthocyanin flowers is 
a possibility, except that glaciata plants are weak, have poor 
branching and  usually not great flowers. But these are not all minor 
problems. There are now stronger and better formed SDB and IB  
glaciatas and eventually perhaps also with TB, but not so far. All TB 
glaciatas I have grown here have quickly disappeared.

The proper test to see if a plant is a glaciata is to cross with a 
plicata. All seedlings will be plicata if it is a glaciata. Of course 
some of thes plicata could be glaciatas.

It actually it is even more complicated, but no further information for 
this post, I sure it is confusing enough as is.

Chuck Chapman

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