Re: squalens, sambucina etc (a bit long) (was Iris pallida cultivars & nomenclatural question)


--- In iris-species@yahoogroups.com, "David Ferguson" 
<manzano57@m...> wrote:
"The original publications of some of these names are as follows:
Iris sambucina Linnaeus 1759, 3A in Systema ed. X:863. 
Iris squalens Linnaeus 1759, 3B in Systema ed. X:863...." [in 
response to a question posed by Linda Mann]

Dave,

I found the information in this post of extraordinary interest, 
especially the extended quote concerning the rules of nomenclature 
for hybrids.  Thank you for posting it.

Just as a comment--could not a case be made for *sambucina* simply 
because it is first--before *squalens*--in Linnaeus' publication in 
1759?  I realize the "first" pertains only to the alphabetical order 
of the names, but with all else equal, it would seem that if a choice 
*must* be made, the occurance of "3A" preceeds "3B" and could be 
argued as having priority?

As a separate issue, the question you raised as to how the "Quaker 
Lady" color type could occur among the typical bicolors, and a white 
occur in progeny from an amoena and *pallida mostrans*--The factors 
involved in the various elements of color involved are not simple.  
There appear to be many steps involved in the synthesis of 
anthocyanins, each presumably controlled by a different enzyme and 
DNA precursor.  A diversion of the path of synthesis of a pigment 
into a different track occurs in a number of cases where white vs. 
violet-blue are concerned.  More than one instance of a white crossed 
with a white, both whites due to genetic recessives, has resulted in 
a violet-blue, simply because the alteration of the synthesis path 
has occurred at a different point in the synthesis.

In other words, there are several different reasons why an iris 
blossom may be white rather than the normal violet-blue.  The white 
ground in an amoena, which I would assume is the same as the white 
ground color in *reginae* or yellow in the typical "variegata" forms 
of *Iris variegata* have a different genetic origin than whites 
appearing among populations of *Iris pallida,* or for that matter, 
those appearing among populations of *Iris aphylla,* as in 'Ostrey 
White.'

In addition, the fall overlay veined pattern or the solid overlay 
pattern, usually associated with veined hafts, is a separate genetic 
issue in amoena, variegata and neglecta bicolors from the color in 
the petal ground.  Linda Mann's suggestion of using the 
term "Umbrata" to describe this overlay is a good one it seems to me, 
and helps underscore the genetic difference between this overlay 
issue from whatever color the ground may be.

Without "Umbrata," but with both yellow and violet-blue present, the 
blossom color resulting would be on the order of the "Quaker Lady" 
blending you mention.

Neil Mogensen   z 7  western NC mountains




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