Re: I. sibirica X I. pseudacorus
- Subject: Re: I. sibirica X I. pseudacorus
- From: d*@aol.com
- Date: Sun, 6 Mar 2011 15:02:18 EST
I'm asuming
you are asking abnout the fertility of the the IB's here. Not the actual
production of one. No one is
actually absolutely sure about why there is fertility in the IB's from such a
cross. Most times the gametes formed are all mixed up. The way the
chromosomes divide up would depend on how much alike the sets are like the
other set, the homology. There is no way to tell that just by looking at
it. That is from experimentation. One plant may tend to
produce mostly SDB type gametes that have a full set of 8 and a full set
of twelve. Others may produce gametes composed of all TB
chromsomes. My experience with the aril quarterbreds has been this. Most of them
are sterile. They are composed of one aril set, 10 (or
11) chromosomes and 3 TB sets. So far all I've been able to get off
of the few seeds I've gotten are halfbreds when crossed to other
halfbreds. You would think that you would get a few quarterbreds but the
seeds either abort or don't survive in the garden for me to see them.
Most quarterbreds are highly infertile with other TB's (Capitola and
Elmohr derived quarterbreds seem to be the exception based on the
Registrations) even though the quarterbreds have three sets of TB
chromosomes. They seem to favor being balanced toward their overall
genetic composition of half aril and half TB (i.e. losing a whole set of aril
chromosomes seems to be a big deal). Producing one
is simple. An SDB contributes a set of 8 and a set of 12.
The TB contributes two sets of 12. Said and done. You have an
IB. That is the foundation of the Class itself.
Example 2: SDB's are amphidiploids that combine the
tetraploid chromosomes of MDB's (32) from pure I. pumila
backgrounds and the 48 chromosomes of the TB's. Each gamete
will get one set of two sets chromosomes and two sets of
twelve for a total of 40 chromosomes. When these plants form
gametes the chromosomes divide the set of 16 I. pumila chromosomes
into two sets of 8 and divide the 24 TB chromosomes into two sets of
twelve. The set of 8 chromosomes pair up and the set of twelve pair
up. The cell divides and each gamete will have one set of eight
and one set of twelve for a total of 20
chromosomes. |
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