HYB: pigments: IMCS X PF bee

Neil wondered offlist about the source of the dark umbrata pattern in
the seedling I posted & in case some of the rest of you are interested &
have some insights, I thought I'd share his thoughts and my reply.

Here's what I've come up with:
1) IMMORTALITY throws mostly white kids for everybody I've talked to.
Second most common is yellow/ cream.
2) only sib from this cross that has bloomed so far was pure pink
3) the IMMXCSONG amoena was one of 3 amoenas/neglectas from around 50
seedlings bloomed from the cross.  2 more were rosy lavender selfs.  The
rest were white to cream.
4) sibling (PFb3) to the pod parent (PFb1) is a showy rose neglecta

Soo, with those clues, my thought is that somehow, these crosses are
reducing the dosage of inhibitors of several pigments at the same time.
And PINK FORMAL doesn't have as many doses of anthocyanin inhibitor as
IMM does.

PFb1 probably has both yellow <and> anthocyanin in both standards and
falls, but more in the falls. I may have done extractions last spring,
but can't find my notes.  I no longer trust my memory, but I <think> the
anthocyanin was a surface wash.  Since it has a sib that is brightly
colored with anthocyanins, tho not as dark as this one, and the IMCS
amoena also had anthocyanin, this one may have just lucked into the
combination that had the least inhibitor of both (or maybe all three? if
water soluble yellow is present) pigments.

I've suspected that the pollen parent of the PINK FORMAL X bee kids was
SPIRIT OF FIJI, but that doesn't answer where the umbrata came from.

Unless the yellows and pinks (PINK FORMAL, SPIRIT OF FIJI) are from I.
variegata background with the anthocyanin overlay hidden by anthocyanin
inhibitor that was somehow partly overcome in the seedlings, and the
rest of the way by CELEBRATION SONG's contribution.

On the other hand, the same pattern, more or less, but more pastel,
showed up in the seedling from IMMORTALITY X PINAFORE PINK.  Color in
both of these seedlings were a surprise.  This one quite a bit more
pleasant <g>

Chuck said that he thought PINAFORE PINK has some anthocyanin pigment -
will check both it and PINK FORMAL for anthocyanin this spring.

Maybe PINK FORMAL has more intensifier hardware than PINAFORE PINK, so
once the pigment can form at all, there is more of it?

Also, AUTUMN SENSATION, in the pedigree of IMMORTALITY, might be umbrata
- it was registered close to the time WHOLE CLOTH came out, so the
"amoena" in pedigree of AS was probably the old "umbrata" type.

AUTUMN SENSATION (G. P. Brown, R. 1961). TB Re 29", Re W4V.   "S flushed
blue fading to pure white; F dark blue-purple. (Snow Carnival X amoena x
Green Dragon) X
    (Fall Fairy x Planet)., Brown's Everblooming Gdn 1962. "

??

<Where did the umbrata (rather conspicuous, as the back of the fall
color, or lack thereof, sure does reveal what's up) come from?
Your description of the Pink Formal baby is more than suggestive.  The
"faint wash" probably is the genetic presence of the
umbrata.  Why it was faint, yet this quite evident is a puzzle.  There
is a possibility that IM conceals something in its makeup, although I
don't know where that would have come from. Neil>
--
Linda Mann east Tennessee USA zone 7/8
East Tennessee Iris Society <http://www.korrnet.org/etis>
American Iris Society web site <http://www.irises.org>
talk archives: <http://www.hort.net/lists/iris-talk/>
photos archives: <http://www.hort.net/lists/iris-photos/>
online R&I <http://www.irisregister.com>

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