----- Original Message -----
From:
n*@charter.net
To: i*@yahoogroups.com
Sent: Saturday, December 04, 2004 12:19
PM
Subject: [iris-photos] Re: PHOTO:TB
Brazen Beauty
Margie, I really don't have a great deal of
experience with plicatas, but from what I've seen, provided the iris is some
kind of plicata, even a single one of the pl loci being the pl(luminata)
variation should be highly visible as a light colored but definite shading
between the veins on both standards and falls, but not occurring in the haft
of the fall and base of the standards.
There was a time when irises like that circulated
on the market. Strong plicata, weak luminata--or in some cases, quite
strong luminata, strong plicata as well. I can't subscript or
superscript (as far as I know) with this window, so can't give the correct
symbols, but perhaps this would be clear enough--
Strong plicata, weak luminata in combination---pl
pl pl pl-lum.
Both plicata and luminata equally expressed with
some strength--pl pl pl-lu pl-lu
Both can be watered down by one or two of
the loci occupied by the allele for glaciata--pl pl-lu pl-a pl-a for example,
giving a pale plicata-luminata, I would think.
Another matter that affects how vivid or dark the
plicata is lies in the locus for the anthocyanin color(s)--it is becoming
quite clear there are different anthocyanins floating around in our modern
gene pool. The anthocyanins show dosage effects, so they can be rich,
medium, pale or absent. How many of the variant anthocyanins can be
manipulated by the plicata set of genes I don't know.
As I understand it, "plicata" genetics can only
work with the anthocyanins already present, however weakly or strongly
made.
The other thing to remember is that when a whole
series of different forms of a gene--alleles--exist, only four normally get
included in any one iris, one for each of the chromosomes that are
fundamentally alike. Pl is non-plicata, pl is normal plicata, as is
illustrated in your photo, pl-lu (however that is symbolized) makes for
luminata, pl-a makes an ice white or yellow or pink--a glaciata. If
any one of the four positions--loci--is occupied by the dominant "normal"
variant (allele) the flower is a self, relative to the plicata
pattern.
What shows in your photo looks to me at first
glance a "normal" plicata--and a beauty at that.
Now, Keith Keppel is the expert among the living
about these matters and he might answer your question quite differently.
He has experience of several decades, a very sharp mind and many tens of
thousands of plicata-line seedlings behind his opinions. His article in
TWOI (pp. 99-107) is a gold mine. Written updated, (which I hope does
happen) it will be a mine producing far more value than mere
gold.
A discussion of plicata genetics can be
found in Kenneth Kidd's article beginning on p.399. That also may
help--or further confuse. Have fun!
Neil Mogensen z 7 western NC
mountains
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