HYB: genetics - line & speckle
iris@hort.net
  • Subject: HYB: genetics - line & speckle
  • From: L* M* <l*@lock-net.com>
  • Date: Fri, 17 Feb 2012 11:53:07 -0500
I hope I don't alienate both Waltah and Chuck by copy this here from FB, but I want to get it into the archives so I can find the info when I forget it!
Walter Moores said: <LINE AND SPECKLE - 'Cinnamon Sun' started the ball rolling on this pattern. It was introduced by Hamner in 1993, and Ghio and Duncan carried the pattern to new heights. I don't have a pic for fear of copyright infringement and don't want to go to jail for stealing one. I hope Chuck Chapman can explain this pattern which is not plicata. If you have pics to add to the discussion, please post them.>
And Chuck Chapman replied: <While many people say that this pattern is not plicata, I probably stand alone in saying that it is. Keppel and Ghio are among many who disagree with me. Ring Around Rosie is a plicata. I have a cross with Ring Around Rosie that gave all plicatas. Spiced Lemon, a cross of Dream of Gold X Cinnamon Sun is a very traditional plicata, The kickers are two genes interacting. One of these is the Ae (enhanced anthocyanin) gene which is the gene from aphylla which gives solid clumps of anthocyanin in the vacuole instead of having anthocyanin just in dissolved form (see my article in AIS Bulletin a few years back) This gene is a dominant. The dominant reduction of anthocyanin gene "I" which can turn a blue iris into a white iris, is only partially dominant with the Ae gene. Thus you have interactions.
So various combinations give odd results.Such as removing anthocyain from rims and most of centre of falls in plicatas inclusion pattern.. Even more so when you combine this with plicata which changes density of plicata distribution. There are inclusion and exclusion genes working with plicata. You don't have anthocyanin in center of petals in classic plicata, this is "exclusion" Sometimes you do, such as with luminata and luminata-plicata. This is inclusion of pattern in center of petals. So when you have inclusion of plicata parent in center of petals combined with Ae and a couple of dosages of "I" you get lines and speckles. Anthocyanin is removed from rims and almost all from center of the falls
Crossing a line and speckles to a plicata gives you an odd mix of patterns, resulting from the recombinations of Ae and "I" genes. Some without any anthocyanin and some solid coloured flowers, which really messes up expected patterns found in plicata crosses without these factors. I have an unpublished article on solid coloured flowers from plicata crosses that I can send to anyone who requests it. That is flowers with four plicata genes, yet are solid coloured.> 

Thanks Chuck (and Walter for asking).
Chuck, I'd like to see a copy of your article. Not sure I'm ready to jump into plicatas, but am close. 

Linda Mann east TN USA zone 7

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