TB -- Mesopotamica Influence


Having been reminded that many
new people have joined the list since
our early discussions about the diploid
to tetraploid conversion, I offer this
brief -- and over-simplified -- summary:

1.  Modern TBs are descended from
both European diploid species and
tetraploids from southwestern Asia, for
details of how this can happen, look for
discussions of the "Triploid Route" in the =

archives.

2.  The first-generation offspring were
thus half European, half Asian, and fully =

fertile.   If line-bred, their advanced =

generations would also be half-and-half.

3. Cold-climate hybridizers backcrossed to
diploids for gardenability and diversity
of patterns and colors, while warm-climate =

hybridizers backcrossed to tetraploids
for size -- and later hybridizers crossed things =

from the cold-climate and warm-climate lines
in hopes of getting the best of both worlds.
So we now have some cultivars that are less =

than half Asian and some that are more than
half Asian, in addition to the half-and-half that
we would expect.

Catch #1 -- the original conversion took =

place so many generations ago that we can't
trace individual cultivars to their original species.

Catch #2 --Although the diploid and tetraploid
species are highly cross-sterile because of =

their differing counts, enough of their individual
chromosomes are homologous that not only
are those rare first-generation offspring fully
fertile but theiradvanced-generation offspring =

function more like autotetraploids than =

amphidiploids. In plain English, today we can
be more concerned with individual genes =

than with chromosome sets.

The proverbial bottom line?  What has all
this to do with I. mesopotamica?  It is, by
reputation, the tenderest of the Asian
tetraploid TBs.  =


Hybridizers who want to minimize its
influence can do soby analyizing pedigrees
and using breeding stock  developed in cold, =

wet climates.  Gardeners with cold and =

wet climates will benefit by choosing
cultivars from breeders in similar climates.  =


If you want to know more, ask questions
or search the archives for species origin
of TBs.

Sharon McAllister
73372.1745@compuserve.com





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