Re: HYB: inheritance of rims?

> From: Linda Mann 
> My favorite patterns are the light tops/dark falls with light rims.
> 
> These mostly seem to come in four types -(1) yellow pigmented ground with
> anthocyanin "shadow" on the falls (variegatas); (2) pink ground with
> anthocyanin on the falls (?not sure what these might be called); (3)
white
> ground with anthocyanin on the falls (amoenas with light rims); and (4)
> anthocyanin ground with more pigment on the falls (neglectas with light
> rims).

Linda,

The classic "Emma Cook" pattern is white standards and falls with a
non-plicata anthocyanin rim on the falls only. According to a recent post
from Fred Kerr, who has done a lot of work with this pattern, it is the
effect of complete homozygosity (4 doses) of the PROGENITOR "Dominant
Amoena" allele. The classic EC pattern occurs in irises which also have
genes for the complete inhibition of carotenoid pigments.

Your patterns #1 and 2 represent the expression of the "Emma Cook" pattern
in irises in which the carotenoid pigments are also expressed. In #1 the
pigment is yellow; in #2 the yellow pigment is replaced by the recessive
pink (lycopene) pigment (if you meant that the anthocyanin is on the rims
of the falls).

If you meant for #2 that the anthocyanin is in the center of the falls with
a ligher, rather than a darker rim, then all pedigrees of irises exhibiting
that kind of pattern that I have looked at, and which can be traced far
enough backward, lead to WINE AND ROSES (D. Hall, R&I 1963). The Checklist
description of W&R is "S. rose-pink; F. violet purple edged rose-pink."
Unfortunately, the only pedigree info on W&R in the Checklist is "from two
sdlgs of same coloring", so the origin of the pattern cannot be traced
further back by that means.

Possibly, your patterns #3 and #4 probably represent the W&R rim in an iris
that has
carotenoid expression suppressed. In #3 there would be full, and in #4
partial,
PROGENITOR Dominant Amoena suppression of anthocyanin in the standards (1
or 2 doses(?)).

There is a third genetically different pattern that produces distinct,
non-plicata bands on the rims of the falls that is not represented in your
four examples. That is the "Joyce Terry" pattern, which affects only
carotenoid pigments and produces flowers with reduced carotenoid diffused
throughout the standards and white falls with a more intrnsely pigmented
carotenoid rim. This might better be called the "Palomino" pattern as the
cultivar of that name (D. Hall, R. 1951, I. 1952) is the first exemplar. It
is apparently a recessive pattern that arose from Hall's tangerine-bearded
pink breeding lines.

There is also presumably still another gene effect responsible for the
light colored fall rim on recessive amoenas/variegatas, such as WABASH. 

Jeff Walters in northern Utah  (USDA Zone 4/5, Sunset Zone 2, AHS Zone 7)
jcwalters@bridgernet.com

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