HYB: genetics - luminata - pl-lu, pl-a, and pl
iris@hort.net
  • Subject: HYB: genetics - luminata - pl-lu, pl-a, and pl
  • From: L* M* <l*@lock-net.com>
  • Date: Sun, 28 Oct 2012 12:55:27 -0400
This is a discussion about luminata genetics currently on FaceBook that started about a photo Ginny Spoon posted. I'm pasting it here so we can all find it again for reference if/when we want to. This is the first part of the discussion:
Chuck Chapman .... pl-a and pl-lu are co-dominant. All luminata patterned iris are a combination of pl-a and pl-lu. My suspicions are that an iris with 4 pl-lu would be solid purple. But we so far have no plants that we can point a finger at, and say that it is 4 pl-lu, and be sure that it is. So thus my high interest in it. Without test crosses to confirm, we can only speculate.
Tom L Waters Chuck, I guess my information is out of date (not surprising, since most of what I know is from my initial study ~30 years ago), but I thought early crosses with luminatas and glaciatas established that pl and pl-lu are codominant, and both dominant with respect to pl-a. Did people later see clear dosage effects for pl-a v. pl-lu?
Chuck Chapman There has been a lot of testing of this by myself, by Don, Keith Keppel, and Christopherson and perhaps others as well. several write ups a well. Crossing luminatas shows that almost always you get glaciatas drop out. Were you don't, there is small numbers in seedlings. Most common genetics in luminata pattern is 3 pl-a, 1 pl-lu. the other combinations can also show luminata pattern, but less of the pattern (most of time). We all have been looking for the 4 pl-lu. From what I can tell pl-a is responsible for the absence of pigment in veins of luminata pattern. So question is, what does pl-lu gene do? I suspect that 4 pl-lu is solid coloured. So basically luminata is a phenotype, not a genotype.
Tom L Waters Thanks Chuck! Let me paraphrase to check that I understand. pl-lu is thus an allele that allows for partial expression of pl-a, but is otherwise probably neutral in effect. Different from PL which completely dominates, and different from pl which gives plicatta. So "fancy plicatas" that display both plicata markings and luminata markings would have to have some combination of pl, pl-lu, and pl-a? 
about an hour ago 7
Tom L Waters 4 x pl-lu would thus be the simplest way to test the ideas. If the, old notion is correct, it should appear as a luminata and produce only luminata seedlings when selfed. If the idea you describe is correct, it should not appear to be a luminata at all, as pl-a is needed to contribute to making the pattern. 
about an hour ago 7
Chuck Chapman Linda- it is OK to repost. Tom Yes. It seems that plicata gene effect is more complicated than older ideas suggest. One main confusing factor, that is usually overlooked, is the Ae (anthocyanin enhancement gene). This is the gene that enhances the depth of colour of anthrocyanin through process of producing solid clumps of anthocyanin in cell vacuole. Known as AVI, or anthrocyanin vascular inclusions. Present in most aphylla cultivars, also in other species. Most modern luminatas also have this gene. I'm not sure if it is needed to produce luminatas. But it does effect appearance of luminata flowers dramatically. Which makes it very hard to sort out effects of pl-a and pl-lu, as Ae is often present. In this genetic combination the effects of Ae on pattern expression is not exactly clear. 
a few seconds ago 7

---------------------------------------------------------------------
To sign-off this list, send email to majordomo@hort.net with the
message text UNSUBSCRIBE IRIS
Other Mailing lists | Author Index | Date Index | Subject Index | Thread Index