Re: HYB: REF: umbrata overlay

The whole matter of amoena/variegata is a combination of traits.

A normal *pallida* blue, when crossed with a diploid amoena, will result in
mostly neglectas.  The fall overlay (Umbrata) is a separate matter from the
non-blue ground color of the amoena or variegata and will appear in many of
the seedlings--I would expect about three out of four on the average.

Neglectas crossed with amoenas will give a combination of amoenas and
neglectas depending on the genetic makeup of the neglecta.  This has to do
with the ground color, not the overlay.

In both diploids and tetraploids, the Umbrata can vary from butterfly-wing
veins patterns, displayed by many species *variegata* clones, the diploid
origin of the Umbrata pattern.  I suspect some of the asiatic tetraploids have
a version of the Umbrata, as a number of them are definitely
bitones--neglectas or bicolor blends.

BRUNO, as one of the early tetraploid bicolors, displays a lot of yellow, some
violet-blue, making the ground color brownish yellow, with a strongly
developed, very solid fall overlay.  I would expect outcrosses from BRUNO to a
yellow to result in some yellows, with a smaller proportion of yellows with
some degree of fall overlay development.

At least two separate factors are at work in an amoena such as WABASH--the
recessive non-blue ground color, plus the dominant fall overlay.

Neil Mogensen

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